[1] DRISKELL R R,CLAVEL C,RENDL M,et al.Hair follicle dermal papilla cells at a glance[J].J Cell Sci,2011,124(8):1179-1182. [2] 谢文燕,王修启,严会超,等.家禽羽毛生长发育规律及其调控机制[J].动物营养学报,2017,29(10):3452-3459. XIE W Y,WANG X Q,YAN H C,et al.Poultry feathers:growth,development and regulation mechanism[J].Chinese Journal of Animal Nutrition,2017,29(10):3452-3459.(in Chinese) [3] SARAGIH H T S G,PERDAMAIAN A B I,SADIMAN,et al.Plumage colours Stability in Inbreed Pelung Chicken[J].BIO Web Conf,2021,33:01005. [4] HAFEZ R D A,HADZIMA-NYARKO M,AHMED S M,et al.Recycled chicken feather sand as a partial replacement for natural sand for producing eco-friendly mortar[J].Buildings,2023,13(2):421. [5] 周泽宇,张夕霏,孟 婕,等.鸡羽色研究现状及发展[J].河北科技师范学院学报,2022,36(3):68-72,80. ZHOU Z Y,ZHANG X F,MENG J,et al.Research status and development of chicken feather color[J].Journal of Hebei Normal University of Science & Technology,2022,36(3):68-72,80.(in Chinese) [6] KANAKACHARI M,ASHWINI R,CHATTERJEE R N,et al.Embryonic transcriptome unravels mechanisms and pathways underlying embryonic development with respect to muscle growth,egg production,and plumage formation in native and broiler chickens[J].Front Genet,2022,13:990849. [7] KAUR S,KUMAR S,KUMAR P.Association between plumage colour and body weight in native chickens[J].J Anim Res,2021, 11(3):527-537. [8] LI R T,WANG X L,WANG Y X,et al.Research note:combined analysis of BSA-Seq based mapping and RNA-Seq reveals candidate genes associated with sub-columbian plumage in h line chickens[J].Poult Sci,2023,102(6):102665. [9] HUA G Y,CHEN J F,WANG J,et al.Genetic basis of chicken plumage color in artificial population of complex epistasis[J].Anim Genet,2021,52(5):656-666. [10] CHO E,KIM M,MANJULA P,et al.A retroviral insertion in the tyrosinase (TYR) gene is associated with the recessive white plumage color in the Yeonsan Ogye chicken[J].J Anim Sci Technol,2021,63(4):751-758. [11] 吴日富,瞿 浩,严 霞,等.家鸡羽色性状遗传调控机制研究进展[J].中国畜牧兽医,2022,49(5):1806-1816. WU R F,QU H,YAN X,et al.Research progress on the genetic regulation mechanism of feather color traits in domestic chickens[J].China Animal Husbandry & Veterinary Medicine,2022,49(5):1806-1816.(in Chinese) [12] FUNK E R,SPELLMAN G M,WINKER K,et al.The genetic basis of plumage coloration and elevation adaptation in a clade of recently diverged alpine and arctic songbirds[J].Evolution,2023,77(3):705-717. [13] SMYTH JR J R,BOISSY R E,FITE K V.The DAM chicken:a model for spontaneous postnatal cutaneous and ocular amelanosis[J].J Hered,1981,72(3):151-156. [14] DANIELS T.Fayoumi chickens[EB/OL].Poultry Keeper,(2015-03-25)[03-21].https://poultrykeeper.com/chicken-breeds/fayoumi-chickens/. [15] DANIELS T.Campine chickens[EB/OL].Poultry Keeper,(2023-03-25)[03-21].https://en.wikipedia.org/wiki/Campine_chicken. [16] WESTFÄLISCHE-TOTLEGER.Westfälische totleger[EB/OL].Jugendseite Westfalen.http://jugendseite-westfalen. de/Rassege fluegel- lexikon/huehner/Westfaelische%20Totleger.html. [17] HATCHERY D.Barred-plymouth-rock[EB/OL].Caldwell:Dunlap Hatchery Poultry.[2015-03-21].https://www.dunlaphatcherypoultry.com/product/barred-plymouth-rock/. [18] THALMANN D S,RING H,SUNDSTRÖM E,et al.The evolution of Sex-linked barring alleles in chickens involves both regulatory and coding changes in CDKN2A[J].PLoS Genet,2017,13(4):e1006665. [19] 符 蓉.利用CDKN2A基因上的SNP标记建立鸡芦花基因的分子检测技术[D].广州:华南农业大学,2019. FU R.Genotyping method for sex-linked barring in chicken with the SNP markers in CDKN2A gene[D].Guangzhou:South China Agricultural University,2019.(in Chinese) [20] SCHWOCHOW D,BORNELÖV S,JIANG T X,et al.The feather pattern autosomal barring in chicken is strongly associated with segregation at the MC1R locus[J].Pigment Cell Melanoma Res,2021,34(6):1015-1028. [21] 何艺平,许继国,谢袖娟,等.鸡羽色性状基因座研究进展[J].广东农业科学,2014,41(9):129-134. HE Y P,XU J G,XIE X J,et al.Research progress of feather color loci in Gallus gallus domestica[J].Guangdong Agricultural Sciences,2014,41(9):129-134.(in Chinese) [22] HELLSTRÖM A R,SUNDSTRÖM E,GUNNARSSON U,et al.Sex-linked barring in chickens is controlled by the CDKN2A/B tumour suppressor locus[J].Pigment Cell Melanoma Res,2010,23(4):521-530. [23] D’ALBA L,KIEFFER L,SHAWKEY M D.Relative contributions of pigments and biophotonic nanostructures to natural color production:a case study in budgerigar (Melopsittacus undulatus) feathers[J].J Exp Biol,2012,215(Pt 8):1272-1277. [24] ROULIN A,DUCREST A L.Genetics of colouration in birds[J].Semin Cell Dev Biol,2013,24(6-7):594-608. [25] CAREFOOT W C.Effect of the eumelanin restrictor Db on plumage pattern phenotypes of the domestic fowl[J].Br Poult Sci,1985,26(3):409-412. [26] ANDERSSON L,BED’HOM B,CHUONG C M,et al.The genetic basis for pigmentation phenotypes in poultry[M]//AGGREY S E,ZHOU H J,TIXIER-BOICHARD M,et al.Advances in Poultry Genetics and Genomics.London:Burleigh Dodds Science Publishing, 2020: 67-106. [27] CAREFOOT W C.Inheritance of the barred plumage pattern of the Silver Campine Fowl,together with its relationship to other patterned fowl[J].Br Poult Sci,1999,40(2):217-220. [28] 聂昌盛,曲鲁江.鸡色素沉积相关性状遗传基础的研究进展[J].中国畜牧杂志,2018,54(6): 6-12. NIE C S, QU L J. Advances in genetic research of pigment traits in chicken[J].Chinese Journal of Animal Science,2018,54(6): 6-12.(in Chinese) [29] PRUM R O,WILLIAMSON S.Reaction-diffusion models of within-feather pigmentation patterning[J].Proc Biol Sci,2002,269(1493):781-792. [30] INABA M,JIANG T X,LIANG Y C,et al.Instructive role of melanocytes during pigment pattern formation of the avian skin[J].Proc Natl Acad Sci U S A,2019,116(14):6884-6890. [31] INABA M,CHUONG C M.Avian pigment pattern formation:developmental control of macro- (across the body) and micro- (within a feather) level of pigment patterns[J].Front Cell Dev Biol,2020,8:620. [32] LIN S J,FOLEY J,JIANG T X,et al.Topology of feather melanocyte progenitor niche allows complex pigment patterns to emerge[J].Science,2013,340(6139):1442-1445. [33] FAN Y K,WU X W,LI Y M,et al.Effect of polymorphisms in the 5'-flanking sequence of MC1R on feather color in Taihang chickens[J].Poult Sci,2022,101(12):102192. [34] YANG C W,RAN J S,YU C L,et al.Polymorphism in MC1R,TYR and ASIP genes in different colored feather chickens[J].3 Biotech,2019,9(5):203. [35] RODRIGUES A R,ALMEIDA H,GOUVEIA A M.Intracellular signaling mechanisms of the melanocortin receptors:current state of the art[J].Cell Mol Life Sci,2015,72(7):1331-1345. [36] SPILLMAN W J.Spurious allelomorphism:results of some recent investigations[J].Amer Nat,1908,42(501):610-615. [37] JAMES B J.Linear relationship of the loci for barring,dermal melanin inhibitor,and recessive white skin on the chicken Z chromosome[J].Poult Sci,1988,67(4):530-533. [38] DORSHORST B J,ASHWELL C M.Genetic mapping of the sex-linked barring gene in the chicken[J].Poult Sci,2009, 88(9): 1811-1817. [39] 席 洋.基于基因组和转录组测序揭示鸭黑色和褐色羽形成的分子机制[D].雅安:四川农业大学,2020. XI Y.Genome and transcriptome sequencing reveal the molecular mechanism of black and brown plumage phenotypes in ducks[D].Yaan:Sichuan Agricultural University,2020.(in Chinese) [40] LI R T,WANG Y X,LIU Y H,et al.Effects of SLC45A2 and GPNMB on melanin deposition based on transcriptome sequencing in chicken feather follicles[J].Animals,2023,13(16):2608. [41] FU Q H,WANG P,ZHANG Y R,et al.Effects of dietary inclusion of asiaticoside on growth performance,lipid metabolism,and gut microbiota in yellow-feathered chickens[J].Animals,2023,13(16):2653. [42] ANDERSSON L.Mutations in domestic animals disrupting or creating pigmentation patterns[J].Front Ecol Evol,2020,8:116. [43] DU W Y,YANG Z L,XIAO C,et al.Identification of genes involved in regulating the development of feathered feet in chicken embryo[J].Poult Sci,2023,102(9):102837. [44] QI Y,ZHANG X,PANG Y,et al.Identification of polymorphism in the MC1R gene and its association with the melanin content in feathers of Chinese yellow quails[J].Braz J Poult Sci,2023,25(2):1-6. [45] JI G G,ZHANG M,TU Y J,et al.Molecular regulatory mechanisms in chicken feather follicle morphogenesis[J].Genes, 2023, 14(8):1646. [46] SQUARE T A.Neural crest cell development and evolution[D].Boulder:University of Colorado,2017. [47] WEHRLE-HALLER B,MELLER M,WESTON J A.Analysis of melanocyte precursors in Nf1 mutants reveals that MGF/KIT signaling promotes directed cell migration independent of its function in cell survival[J].Dev Biol,2001,232(2):471-483. [48] LIN J Y,FISHER D E.Melanocyte biology and skin pigmentation[J].Nature,2007,445(7130):843-850. [49] FAAS L,ROVASIO R A.Distribution patterns of neural-crest-derived melanocyte precursor cells in the quail embryo[J].Anat Rec,1998,251(2):200-206. [50] GUNNARSSON U,KERJE S,BED’HOM B,et al.The Dark brown plumage color in chickens is caused by an 8.3-kb deletion upstream of SOX10[J].Pigment Cell Melanoma Res,2011,24(2):268-274. [51] KIM S H,MITCHELL M,FUJII H,et al.Absence of p16INK4a and truncation of ARF tumor suppressors in chickens[J].Proc Natl Acad Sci U S A,2003,100(1):211-216. [52] HAUPT Y,MAYA R,KAZAZ A,et al.Mdm2 promotes the rapid degradation of p53[J].Nature,1997,387(6630):296-299. [53] SHARPLESS N E.INK4a/ARF:A multifunctional tumor suppressor locus[J].Mutat Res,2005,576(1-2):22-38. [54] 周成浩,殷建玫,李国辉,等.基于简化基因组测序技术的汶上芦花鸡选择信号分析[J].农业生物技术学报,2022,30(12): 2385-2395. ZHOU C H,YIN J M,LI G H,et al.Selection signal analysis of Wenshang barred chicken (Gallus gallus) based on RAD-seq technology[J].J Agric Biotechnol,2022,30(12):2385-2395.(in Chinese) [55] KOBAYASHI T,URABE K,WINDER A,et al.Tyrosinase related protein 1 (Trp1) functions as a DHICA oxidase in melanin biosynthesis[J].EMBO J,1994,13(24):5818-5825. [56] NAM I S,OH M G,NAM M S,et al.Specific mutations in the genes of MC1R and TYR have an important influence on the determination of pheomelanin pigmentation in Korean native chickens[J].J Adv Vet Anim Res,2021,8(2):266-273. [57] LE L,ESCOBAR I E,HO T,et al.SLC45A2 protein stability and regulation of melanosome pH determine melanocyte pigmentation[J].Mol Biol Cell,2020,31(24):2687-2702. [58] 张 静.鹅羽色相关候选基因ASTP和TYRP1基因克隆、表达及SNP检测[D].重庆:西南大学,2015. ZHANG J.Molecular Clonging,expression and SNP detection on ASIP andTYRP1 in the candidate genes of goose plumage color[D].Chongqing:Southwest University,2015.(in Chinese) [59] 宋素芳,宋幸辉,蔺成刚,等.SLC45A2基因在“豫粉1号”浅芦花羽色中的表达及酪氨酸对其在黑色素细胞中表达量的影响[J].中国畜牧杂志,2019,55(9):64-67. SONG S F,SONG X H,LIN C G,et al.Expression of SLC45A2 in columbian plume color of "Yufen Ⅰ" chicken and effect of tyrosine on its expression in melanocytes[J].Chinese Journal of Animal Science,2019,55(9):64-67.(in Chinese) [60] 刘小辉.坝上长尾鸡资源保护及羽色多样性分子遗传基础[D].保定:河北农业大学,2018. LIU X H.Resources protection and molecular genetic basis of plumage color diversity of Bashang long-tail chickens[D].Baoding: Hebei Agricultural University,2018.(in Chinese) [61] 许继国,张细权.鸡芦花羽色性状的遗传和应用[J].养禽与禽病防治,2011(8):4-5. XU J G,ZHANG X Q.Heredity and application of feather color traits in Luhua chicken[J].Poultry Husbandry and Disease Control,2011(8):4-5.(in Chinese) [62] WANG H E,WEN J H,LI H Y,et al.Candidate pigmentation genes related to feather color variation in an indigenous chicken breed revealed by whole genome data[J].Front Genet,2022,13:985228. [63] 杨晓燕.乌骨鸡肤色遗传研究[D].北京:中国农业大学,2004. YANG X Y.Genetic research on skin color of black-bone chicken[D].Beijing:China Agricultural University,2004.(in Chinese) [64] ALESSIO B M,GUPTA A.Diffusiophoresis-enhanced Turing patterns[J].Sci Adv,2023,9(45):eadj2457. |